-:
$%&*5 J/9%&*5 -1$#5 =>7% &+"$*, ,:/<@(5, =4>"(µ# µ7 +$#>"3,7%5. L% ,$#:9+%*-7"75 &+??%7"4#1µ7/75
$#%&%?875 +/<&#:/ ,-#/ -1$# +:-*. L -:$%&*5 -1$#5 -.5 J/9#&8/+5 &+% O%?%$$%/3/ =>7% &+"$#15 µ7 >"3µ+
(
>"* $"',%/# $"#5 &%-"%/($*, µ7 ?8475 < &+@*?#: +$#>"3,7%5. !+ )"#1-+ +:-#1 -#: -1$#: 78/+% >("85
8/75, 4?:&' &
+% $?#1,%+ ,7 '"(µ+. 27/%&', -+ 9=/-"+ +:-' 78/+% $%# $+"+4(4%&' +$* 7&78/+ -#: J/9%&#1
-1
$#:, +??' 7$7%9< -+ )"#1-+ -#:5 97/ )="#:/ >"(µ+-%,µ*, 97/ $"#-%µ#1/-+% +$* -#:5 &+-+/+?(-=5.
G7 E',. -#:5 9:# $%# $'/( -1$#:5, =>#:/ 9.µ%#:"4.@78 /=75 $#%&%?875, *$(5 . J"#:8/, !*µ% F-&%/5, 07/-,
P=$-+>, C#:85, D8"% &+% '??75.
I 0#:'E+ (
Psidium guajava
-#%&#4
=/7%+
Myrtaceae
), 78/
+% 9=/-"# +7%@+?=5, @+µ/3975 &+% %@+47/=5 -(/
-"#
$%&3/ $7"%#>3/ -.5 6µ7"%&<5. Q>7% &#"µ* ?78# µ7 )?#%* $#: K7)?#:98A7-+% ,7 ?7$-' ?=$%+. L% &+"$#8
-.5 :
$'4#/-+% ,-./ &+-.4#"8+ µ#1"(/ &+% =>#:/ 9%'µ7-"# +$* 4-12 7&. Q>#:/ ?7$-< )?#19+ &+% -# 7,(-
-7"%&* -#:5 78/
+% ,+"&3975 µ7 &=/-"# $#?-3975, 47µ'-# ,$="µ+-+. I 0#:'E+ 7:9#&%µ78 ,7 µ74'?# 71"#5
79
+)%&3/ ,:/@.&3/, (,-*,# $"#-%µ' 4*/%µ+ &+% $?#1,%+ ,7 #"-
4
+/%&< #:,8+ 79').. 6/-=>7% ,7 µ+&"=5 $7"%*9#:5 79+)%-
&
<5 K."+,8+5, +??' +/=>7-+% &+% µ+&"=5 $7"%*9#:5
:
$7"E#?%&<5 79+)%&<5 :4"+,8+5. R,-*,#, 4%+ &+?<
$+"+4(4< &+% $#%*-.-+ )"#1-(/, -+ 9=/-"+ 97/
$"=$7% /+ 9%H',#:/, #1-7 /+ -#:5 98/7-+% $7"%,-
,*-7".
$#,*-.-+ /7"#1 +$* +:-</ $#: >"7%-
'
A#/-+%. R5 -"#$%&* 9=/-"# 7$."7'A7-+% +$* -#
H1>#5, (,-*,# +/-=>7% $7"%,,*-7"# +$* '??+
-"#
$%&' 789. &+%, ,-./ 01$"#, µ$#"78 /+ &+??%7"-
4.@78 &
+% ,7 $79%/=5 $7"%#>=5 $="+/ -.5 $+"+?%+-
&
<5 A3/.5. L% >+µ.?=5 @7"µ#&"+,875 K."+8/#:/ &+%
-#:5 &?'9#:5 -#: 9
=/9"#:, (,-*,# +:-* E4'A7% 4"<-
4#"
+ /=+ E?',-.,. µ7-' -./ $+"=?7:,< -#:5. I 0#:'E+
$#??+$?+,%'A7-+%, &:"8(5, µ7 ,$*"#. L% ,$*"#% $+8"/#/-+%
µ*/# +$* 9=/-"+ $#: $+"'4#:/ &+?<5 $#%*-.-+5 )"#1-+ &+% )="#:/
+:-#4*/%µ+ '/@., =-,% 3,-7 /+ µ7%3/7-+% # &8/9:/#5 $+"+4(4<5 97/9":??8(/ +µ)8E#?.5 +K8+5. !+ 97/-
9"1??%
+ $#: $+"'4#/-+% +$* ,$*"#:5 7$%?74µ=/(/ 9=/-"(/, 97/ >"7%'A#/-+% 7µE#?%+,µ*. L% ,$*"#% E?+-
,-'/#:/ ,7 9%',-.
µ+ 2-3 E9#µ'9(/. B7 $7"8$-(,. $#: 7)+"µ#,-78 7µE#?%+,µ*5 ,7 ,$#"*):-#, +:-*5
48/7-
+% µ7 -./ -7>/%&< -#: 74&7/-"%,µ#1 $"',%/#: K1?#: &+% µ$#"78 /+ 48/7% #$#%+9<$#-7 $7"8#9# -#:
>"*/#:, )-'/7% -# :$#&78µ7/# /+ =>7% -#:?'>%,-# 5 >%?%#,-' 9%'µ7-"#. S$'">#:/ 9:# -1$#% 0#:'E+5, #%
&%-"%/*,
+"&75 &+% #% "#9*,+"&75. L% &%-"%/*,+"&75, &+-' -./ ("8µ+/,., =>#:/ >"3µ+ 7?+)"35 &8-"%/#
-*,# 7
K(-7"%&', *,# &+% 7,(-7"%&'. Q>#:/ 471,. 4?:&*K%/. $#: $?.,%'A7% -. 471,. -#: 3"%µ#: ?7µ#/%#1,
4%
+ +:-* &+% #/#µ'A#/-+% ;=µ#/ 0#1+E75. L% "#9*,+"&75 $#%&%?875, &+-' -./ ("8µ+/,., =>#:/ >"3µ+ "#-
9*
>"#:/ 7,(-7"%&', +??' &+% µ7"%&35 7K(-7"%&'. B7 +/-8@7,. µ7 -./ &%-"%/*,+"&. 0#:'E+, . 471,. -.5
4?:&8
A7%, *$(5 -.5 )"'#:?+5. L% 0#:'E75 &+-+/+?3/#/-+% )"=,&%75 < >".,%µ#$#%#1/-+% 4%+ $+"+4(4<
µ+"µ7?'9(/, >:µ3/ &+% $+4(-3/.
I B%7"%µ*4%+ (
Annona cherimolia
-#%&#4
=/7%+
Annonaceae
) 78/
+% 7/9.µ%&* 789#5 -(/ F/97(/, 7/3, ,<µ7"+,
&
+??%7"478-+% ,7 $#??=5 $7"%#>=5 µ7 :$#-"#$%&* &?8µ+, &+@35 &+% ,7 :H8$79+ -.5 -"#$%&<5 A3/.5. D"*-
&7%-
+% 4%+ ):??#E*?# 9=/-"#, -# #$#8# 7&-*5 +$* ):??*$-(,., &+-' -. >7%µ7"%/< $7"8#9#, $+"#:,%'A7%
K."'/,7%5 /7+"3/ E?+,-3/ &+% µ%&"3/ &?'9(/, $+"#:,%'A#/-+5 =-,% µ%+ +,@7/%&< 7µ)'/%,. µ=>"% /+ 7µ-
)
+/%,-#1/ -+ /=+ )1??+ &+% '/@. &+-' -./ F/#%K.. !+ '/@. -.5 B%7"%µ*4%+ 78/+% 7"µ+)"*9%-+, (,-*,# $+-
"#:,%'
A#:/ =/+ µ.>+/%,µ* µ7 -#/ #$#8# +$#)7147-+% . +:-#4#/%µ#$#8.,.. B:4&7&"%µ=/+, -+ '/@. -+
#
$#8+ 78/+% E"+>1E%+, -%5 $"(%/=5 3"75 ,:µ$7"%)="#/-+% (5 @.?:&', 7/3, ,-+9%+&', 7K7?8,,#/-+% ,7 +"-
,7/%&'
µ=>"% -# +$*47:µ+ -.5 89%+5 µ="+5. C$#µ=/(5, 4%+ /+ 7K+,)+?%,-78 . $+"+4(4< 78/+% +$+"+8-.-.
. ,-
+:"#4#/#µ#$#8.,.. 6:-* 7$%-:4>'/7-+% µ7 ,:4&+??%="47%+ ,:/+@#:,3/ &+% ,:µE+-3/ $#%&%?%3/,
&
+@35 &+% µ7 -7>/.-< 4#/%µ#$#8.,.. L &+"$*5 78/+% ,:4&'"$%# &+% -# ,><µ+ -#: $#%&8??7% +$* &(/%&* =(5
(#7%9
=5. I 7$%)'/7%+ -#: &+"$#1 78/+% $"',%/. &+% ?7$-<, 7/3 7µ)+/8A7% $"#7K#>=5 ,$7%"#7%935 9%+-7-
-
+4µ=/75. I ,'"&+ 78/+% &"7µ ?7:&<, µ+?+&<, 47:,-%&<, +"(µ+-%&<, 7?+)"' :$*K%/. &+% $7"%=>7% $#??#15
!"#
$%&'
!
7